| Name | Hold | Shares | Value | Type |
|---|---|---|---|---|
| Flagship Pioneering Inc | 22.41% | 12,674,120 | $56.78M | Institution |
| Fmr LLC | 9.68% | 5,474,491 | $24.53M | Institution |
| Bvf Inc | 9.38% | 5,302,000 | $23.75M | Institution |
| Cigall Kadoch | 6.41% | 3,624,379 | $16.24M | Insider |
| Deerfield Management Company LP | 5.76% | 3,255,514 | $14.58M | Institution |
| Raymond James Financial Inc | 4.35% | 2,462,488 | $11.03M | Institution |
| Blackrock Inc | 4.00% | 2,259,948 | $10.12M | Institution |
| Vanguard Group Inc | 3.36% | 1,899,457 | $8.51M | Institution |
| Flagship Ventures Fund V General Partner LLC | 3.27% | 1,851,801 | $8.30M | Insider |
| Euclidean Capital LLC | 2.78% | 1,573,761 | $7.05M | Institution |