| Name | Hold | Shares | Value | Type |
|---|---|---|---|---|
| Flagship Pioneering Inc | 22.42% | 12,674,120 | $51.58M | Institution |
| Fmr LLC | 9.79% | 5,534,516 | $22.53M | Institution |
| Bvf Inc | 9.38% | 5,302,000 | $21.58M | Institution |
| Cigall Kadoch | 6.41% | 3,624,379 | $14.75M | Insider |
| Deerfield Management Company LP | 5.76% | 3,255,514 | $13.25M | Institution |
| Raymond James Financial Inc | 4.19% | 2,367,292 | $9.63M | Institution |
| Blackrock Inc | 3.94% | 2,229,133 | $9.07M | Institution |
| Vanguard Group Inc | 3.33% | 1,884,673 | $7.67M | Institution |
| Flagship Ventures Fund V General Partner LLC | 3.28% | 1,851,801 | $7.54M | Insider |
| Euclidean Capital LLC | 2.78% | 1,573,761 | $6.41M | Institution |