| Name | Hold | Shares | Value | Type |
|---|---|---|---|---|
| Flagship Pioneering LLC | 21.59% | 12,674,120 | $76.93M | Institution |
| Fmr LLC | 10.30% | 6,047,198 | $36.71M | Institution |
| Bvf Inc | 9.95% | 5,842,000 | $35.46M | Institution |
| Deerfield Management Company LP | 7.48% | 4,394,367 | $26.67M | Institution |
| Cigall Kadoch | 6.17% | 3,624,379 | $22.00M | Insider |
| Raymond James Financial Inc | 4.43% | 2,602,382 | $15.80M | Institution |
| Blackrock Inc | 4.03% | 2,364,866 | $14.35M | Institution |
| Flagship Ventures Fund V General Partner LLC | 3.15% | 1,851,801 | $11.24M | Insider |
| Euclidean Capital LLC | 2.68% | 1,573,761 | $9.55M | Institution |
| Vanguard Capital Management LLC | 2.55% | 1,495,089 | $9.08M | Institution |