| Name | Hold | Shares | Value | Type |
|---|---|---|---|---|
| Flagship Pioneering Inc | 22.41% | 12,674,120 | $68.44M | Institution |
| Fmr LLC | 9.68% | 5,474,491 | $29.56M | Institution |
| Bvf Inc | 9.38% | 5,302,000 | $28.63M | Institution |
| Cigall Kadoch | 6.41% | 3,624,379 | $19.57M | Insider |
| Deerfield Management Company LP | 5.76% | 3,255,514 | $17.58M | Institution |
| Raymond James Financial Inc | 4.35% | 2,462,488 | $13.30M | Institution |
| Blackrock Inc | 4.00% | 2,259,948 | $12.20M | Institution |
| Vanguard Group Inc | 3.36% | 1,899,457 | $10.26M | Institution |
| Flagship Ventures Fund V General Partner LLC | 3.27% | 1,851,801 | $10.00M | Insider |
| Euclidean Capital LLC | 2.78% | 1,573,761 | $8.50M | Institution |