| Name | Hold | Shares | Value | Type |
|---|---|---|---|---|
| Flagship Pioneering LLC | 21.59% | 12,674,120 | $56.27M | Institution |
| Fmr LLC | 9.19% | 5,396,814 | $23.96M | Institution |
| Bvf Inc | 9.03% | 5,302,000 | $23.54M | Institution |
| Deerfield Management Company LP | 6.22% | 3,649,211 | $16.20M | Institution |
| Cigall Kadoch | 6.17% | 3,624,379 | $16.09M | Insider |
| Raymond James Financial Inc | 4.27% | 2,505,821 | $11.13M | Institution |
| Blackrock Inc | 4.00% | 2,349,870 | $10.43M | Institution |
| Vanguard Group Inc | 3.29% | 1,928,992 | $8.56M | Institution |
| Flagship Ventures Fund V General Partner LLC | 3.15% | 1,851,801 | $8.22M | Insider |
| Euclidean Capital LLC | 2.68% | 1,573,761 | $6.99M | Institution |