| Name | Hold | Shares | Value | Type |
|---|---|---|---|---|
| Flagship Pioneering LLC | 22.41% | 12,674,120 | $60.84M | Institution |
| Fmr LLC | 9.54% | 5,396,814 | $25.90M | Institution |
| Bvf Inc | 9.38% | 5,302,000 | $25.45M | Institution |
| Deerfield Management Company LP | 6.45% | 3,649,211 | $17.52M | Institution |
| Cigall Kadoch | 6.41% | 3,624,379 | $17.40M | Insider |
| Raymond James Financial Inc | 4.43% | 2,505,821 | $12.03M | Institution |
| Blackrock Inc | 4.16% | 2,349,870 | $11.28M | Institution |
| Vanguard Group Inc | 3.41% | 1,928,992 | $9.26M | Institution |
| Flagship Ventures Fund V General Partner LLC | 3.27% | 1,851,801 | $8.89M | Insider |
| Euclidean Capital LLC | 2.78% | 1,573,761 | $7.55M | Institution |