| Name | Hold | Shares | Value | Type |
|---|---|---|---|---|
| Flagship Pioneering Inc | 22.41% | 12,674,120 | $56.27M | Institution |
| Fmr LLC | 9.68% | 5,474,491 | $24.31M | Institution |
| Bvf Inc | 9.38% | 5,302,000 | $23.54M | Institution |
| Cigall Kadoch | 6.41% | 3,624,379 | $16.09M | Insider |
| Deerfield Management Company LP | 5.76% | 3,255,514 | $14.45M | Institution |
| Raymond James Financial Inc | 4.35% | 2,462,488 | $10.93M | Institution |
| Blackrock Inc | 4.00% | 2,259,948 | $10.03M | Institution |
| Vanguard Group Inc | 3.36% | 1,899,457 | $8.43M | Institution |
| Flagship Ventures Fund V General Partner LLC | 3.27% | 1,851,801 | $8.22M | Insider |
| Euclidean Capital LLC | 2.78% | 1,573,761 | $6.99M | Institution |