| Name | Hold | Shares | Value | Type |
|---|---|---|---|---|
| Flagship Pioneering LLC | 21.59% | 12,674,120 | $52.98M | Institution |
| Fmr LLC | 10.30% | 6,047,198 | $25.28M | Institution |
| Bvf Inc | 9.95% | 5,842,000 | $24.42M | Institution |
| Deerfield Management Company LP | 7.48% | 4,394,367 | $18.37M | Institution |
| Cigall Kadoch | 6.17% | 3,624,379 | $15.15M | Insider |
| Raymond James Financial Inc | 4.43% | 2,602,382 | $10.88M | Institution |
| Blackrock Inc | 4.03% | 2,364,866 | $9.89M | Institution |
| Flagship Ventures Fund V General Partner LLC | 3.15% | 1,851,801 | $7.74M | Insider |
| Euclidean Capital LLC | 2.68% | 1,573,761 | $6.58M | Institution |
| Vanguard Capital Management LLC | 2.55% | 1,495,089 | $6.25M | Institution |